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But seeds aren't photosynthesizing. On the flip side, roots need oxygen or they will rot. But then, seeds aren't rooting.Plants "breath in" co2 during photosynthesis.
According to your link, seeds are about the toughest life form around next to the cockroach.
If sufficiently dried, all of these seeds are effectively dormant to the point that the surrounding gas mix, or lack thereof, is insignificant to storage. Note, too, that we are talking viability, here--the ability for the seed to grow after storage. Seeds do respire at an extremely slow rate, however, which can be measured with a small increase in carbon dioxide gas after many years of storage.
Like I said, life is a wonderment for sure